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📖Breeding Terminology: Essential Glossary for Exams

A comprehensive glossary of exam-critical breeding and genetics terms — heterosis, inbreeding depression, ideotype, chimera, amphidiploid, and more — with precise definitions.

Why Terminology Matters

In competitive exams (IBPS AFO, NABARD, ICAR JRF, RRB SO), one-liner questions on breeding terminology are among the highest-frequency question types. Knowing the precise definitions — and distinguishing between similar-sounding terms like heterosis vs. luxuriance, or synthetic vs. composite — can make the difference between a correct and incorrect answer.


A comprehensive glossary of exam-critical breeding and genetics terms frequently asked in IBPS-AFO, NABARD, ICAR-JRF, and RRB-SO examinations. Each term includes its precise definition and exam context.


Pollination & Reproduction Terms

  • Duodichogamy — individual plants produce two batches of male flowers temporally separated by female flowers. Rarest temporal sexual system in flowering plants. (Asked in RRB SO 2019)

  • Geitonogamy — pollen from one flower of a plant falls on the stigma of another flower of the same plant. Functionally cross-pollination but genetically self-pollination. E.g. Maize.

  • Xenogamy — transfer of pollen from a flower of one plant to the stigma of a flower of a different plant. This is true cross-pollination and leads to maximum genetic recombination.

  • Emasculation — removal of stamens or anthers, or killing of pollen grains, to prevent self-pollination. Essential step in hybridization of bisexual flowers. Methods include hand emasculation, hot water treatment, alcohol treatment, and use of gametocides. (AFO-2023)

  • Parthenocarpy — development of fruit without fertilization. The resulting fruits are seedless. E.g. Banana, Seedless watermelon.

  • Agamospermy — development of embryo and seed without meiosis and fertilization. A form of apomixis where seeds are produced asexually. E.g. Citrus, Mango.

  • Apomixis — reproduction through seeds formed without fusion of gametes (without sexual union). It includes agamospermy, adventitious embryony, and apospory.

NOTE

Geitonogamy vs Xenogamy: Geitonogamy involves pollen transfer between flowers of the same plant (genetically = selfing), while Xenogamy involves pollen transfer between different plants (true outcrossing). This distinction is frequently tested.


Genetic Variation Terms

  • Allosyndesis — pairing of non-homologous chromosomes from diverse sets of an allopolyploid at meiosis. Results in recombination between genomes of different species. (RRB SO-2019)

  • Autosyndesis — pairing of homologous chromosomes from similar sets of an allopolyploid at meiosis. Results in recombination within genomes of the same species.

  • Genepool — sum total of all alleles present in the breeding/reproductive members of a random mating population. Classified into:

    • Primary genepool — same species, cross easily
    • Secondary genepool — related species, cross with some difficulty
    • Tertiary genepool — distantly related, cross with extreme difficulty (embryo rescue needed)
  • Gene frequency — proportion of one allele relative to all alleles of a gene in a Mendelian population. Also called allele frequency.

  • Biodiversity — total variability present within and among species of all living organisms. Includes genetic, species, and ecosystem diversity.

  • Epistasis — when one gene affects the expression of another non-allelic gene. Types include dominant epistasis (12:3:1), recessive epistasis (9:3:4), duplicate epistasis (15:1), complementary epistasis (9:7), and inhibitory epistasis (13:3).

  • Pleiotropism — one gene affects more than one character (pleiotropic genes). E.g. the gene for seed coat colour in peas also affects flower colour and leaf axil pigmentation.

  • Silent Mutation — a mutation that changes the nucleotide sequence but does not alter the amino acid or protein activity due to the degeneracy of the genetic code.

  • Spontaneous Mutations — mutations occurring under natural conditions without any deliberate treatment with mutagens. Frequency is very low (10-5 to 10-8 per gene per generation).

  • Tautomerism — ability of a molecule (especially bases of DNA) to exist in more than one chemical form. Tautomeric shifts in bases lead to transition mutations during DNA replication.

  • Pseudo alleles — genes that govern different expressions of the same character, are closely linked, and exhibit cis-trans position effect. They can be separated by crossing over. E.g. Star and asteroid loci in Drosophila.

IMPORTANT

Allosyndesis vs Autosyndesis is a classic one-liner in RRB SO and AFO exams. Remember: Allo = different (non-homologous pairing), Auto = same (homologous pairing). Both occur in allopolyploids during meiosis.


Breeding & Selection Terms

  • Heterosis (Hybrid Vigour) — superiority of F1 hybrid in one or more characters over its parents. Term coined by G.H. Shull (1914). Three types:

    • Relative heterosis — F1 superiority over mid-parent value
    • Heterobeltiosis — F1 superiority over the better parent
    • Standard/Economic heterosis — F1 superiority over the best commercial check variety
  • Heterobeltiosis — heterosis estimated over the superior/better parent. More useful than mid-parent heterosis for practical breeding purposes.

  • Luxuriance — superiority of F1 over parents in vegetative growth but NOT in yield and adaptation. Also known as pseudo heterosis. The hybrid may look vigorous but performs poorly in reproductive traits.

  • Top cross — cross between an inbred line and an open-pollinated variety. Also known as inbred-variety cross. Used to evaluate general combining ability (GCA) of inbred lines. Proposed by Davis (1927).

  • Inbreeding — mating between individuals related by descent or common ancestry (brother-sister/sib mating). Highest degree of inbreeding is achieved by selfing. Increases homozygosity by 50% per generation in self-pollinated crops.

  • Inbreeding Depression — severe reduction in vigour and fertility in cross-pollinated species when subjected to continued selfing. Very low in self-pollinated crops. Highest in cross-pollinated crops like Carrot, Alfalfa, Lucerne. Opposite of heterosis.

WARNING

Do not confuse Heterosis with Luxuriance. Heterosis shows superiority in yield and economically important traits, while Luxuriance (pseudo heterosis) shows superiority only in vegetative growth without any yield advantage.

  • Acclimatization — adaptation or adjustment of an introduced variety to a new environment. Different from adaptation, which is a natural evolutionary process.

  • Synthetic Variety — produced by crossing in all combinations a number of lines that combine well, followed by open pollination. Can be grown for 4-5 years without significant yield reduction. Reconstituted from parental lines when yield drops. Proposed by Sewall Wright.

  • Composite Variety — produced by mixing seeds of several phenotypically outstanding lines and encouraging open pollination. Broader genetic base than synthetic varieties but may show faster yield decline over generations.

  • Land races — crop cultivars developed through traditional farming over centuries of natural and human selection. Good source of genetic variability and adaptation genes. Also called desi varieties or primitive cultivars.

  • Obsolete varieties — varieties developed by systematic modern agricultural science that have been replaced by newer, improved varieties. Valuable as sources of specific genes.

  • Chimera — an individual with one genotype in some parts and another genotype in other parts. Arises from mutation in a single cell during development. Jaganath rice = gamma-ray induced semi-dwarf mutant from T-141. Types include sectorial, mericlinal, and periclinal chimeras.

  • Amphidiploid — an allopolyploid with two complete copies of each parental genome, behaves as a diploid during meiosis (showing bivalent pairing). Used as a bridge species for character transfer between incompatible species. E.g. Raphanobrassica (2n = 36) — first artificial amphidiploid produced by Karpechenko (1927).

  • Ideotype — according to Donald (1968), a biological model that is expected to perform or behave in a predictable manner within a defined environment. Ideotype breeding aims to combine specific morphological and physiological traits for maximum yield.

TIP

Synthetic vs Composite Variety: Both are used in cross-pollinated crops. Synthetic varieties involve controlled crosses among selected lines (all possible combinations), while Composite varieties are simply mixtures of seeds from outstanding lines. Synthetic varieties maintain yield longer (4-5 years).


Cytogenetics & Molecular Terms

  • Cytogenetics — study of various aspects of chromosomes and their effects on the development of characters. Bridges the gap between cytology and genetics.

  • Genomics — study of the structure and function of the entire genome of an organism. Includes structural genomics, functional genomics, and comparative genomics.

  • Genome — complete set of chromosomes of a diploid species (one haploid set). Represented by letters (e.g. Wheat: AABBDD = 3 genomes).

  • Euphenics — deals with the control of hereditary diseases, especially inborn errors of metabolism, through environmental manipulation or medical intervention. Term coined by Lederberg.

  • Allele — alternate form of a gene occupying the same locus on homologous chromosomes. Shorter term for allelomorph.

  • Multiple alleles — the presence of many alleles of a single gene in a population (but only two per diploid individual). E.g. ABO blood group system has three alleles: IA, IB, i.

  • Antibody — a protein (immunoglobulin) found in serum or plasma, demonstrated by specific reaction with its corresponding antigen. Produced by B-lymphocytes.

  • Antigen — a substance that induces production of a specific antibody when introduced into a vertebrate body. Located on the surface of RBC (in blood group systems).

  • Cytoplasmic inheritance — characters controlled by plasmagenes (genes in cytoplasm — mitochondria, chloroplasts). Also called extranuclear or maternal inheritance. Always shows maternal effect. E.g. Male sterility in maize, Kappa particles in Paramecium.

NOTE

Cytoplasmic inheritance is identified by the fact that reciprocal crosses give different results, and the trait always follows the maternal parent. This is because cytoplasm (and organelles) are inherited almost exclusively from the egg cell.


Important Historical Facts

  • First artificial hybrid in plants: Thomas Fairchild (1717) — crossed Sweet William (Dianthus barbatus) with Carnation (Dianthus caryophyllus). The resulting hybrid is known as Fairchild’s Mule.

  • Back cross — crossing F1 with one of its parents. Important in development of disease resistance through backcross breeding. Two types:

    • Recurrent parent — the parent to which repeated backcrosses are made
    • Donor parent — the parent that donates the gene of interest
  • Test cross — crossing F1 (or any dominant phenotype individual) with its homozygous recessive parent. Used to determine the genotype of a dominant phenotype individual. Proposed by Mendel.

  • Pure line theory — given by Johannsen (1903). A pure line is the progeny of a single homozygous self-pollinated plant. Selection within a pure line is ineffective.

  • Law of Homologous Series — given by N.I. Vavilov (1920). Species that are genetically related exhibit parallel variation in their characteristics.

  • Mutation theory — given by Hugo de Vries (1901) based on his work on Oenothera lamarckiana (Evening Primrose). However, what he observed were actually chromosomal aberrations, not gene mutations.


Quick Revision Table

TermKey PointExam Relevance
DuodichogamyRarest temporal sexual systemRRB SO 2019
AllosyndesisNon-homologous pairing in allopolyploidRRB SO 2019
EmasculationPrevents self-pollination in bisexual flowersAFO 2023
HeterobeltiosisHeterosis over better parentIBPS AFO, NABARD
LuxuriancePseudo heterosis (vegetative only)ICAR JRF
Synthetic varietyLasts 4-5 years without yield dropIBPS AFO
AmphidiploidBridge species, diploid behaviourICAR JRF
IdeotypeDonald (1968), biological modelNABARD
Top crossTests GCA, Davis (1927)AFO
Fairchild (1717)First artificial plant hybridAll exams

TIP

Exam Strategy: Terms like Heterobeltiosis, Luxuriance, Allosyndesis, and Duodichogamy are frequently asked in one-liner format. Memorize the precise definitions and the scientists/years associated with them. Focus on distinguishing between similar-sounding terms (e.g., Allosyndesis vs Autosyndesis, Heterosis vs Luxuriance, Synthetic vs Composite).

Summary Cheat Sheet

Concept / TopicKey Details / Explanation
DuodichogamyTwo batches of male flowers separated by female flowers; rarest temporal sexual system; asked in RRB SO 2019
GeitonogamyPollen from one flower to another flower of the same plant; genetically = selfing (e.g., Maize)
XenogamyPollen from one plant to a different plant; true cross-pollination; maximum genetic recombination
EmasculationRemoval of anthers/stamens to prevent self-pollination; methods: hand, hot water, alcohol, gametocides; AFO-2023
ParthenocarpyFruit development without fertilization; results in seedless fruit (e.g., Banana, Seedless watermelon)
AgamospermySeed and embryo formed without meiosis and fertilization (e.g., Citrus, Mango)
ApomixisReproduction via seeds without gamete fusion; includes agamospermy, adventitious embryony, apospory
AllosyndesisPairing of non-homologous chromosomes in allopolyploids at meiosis; Allo = different; RRB SO 2019
AutosyndesisPairing of homologous chromosomes in allopolyploids at meiosis; Auto = same
GenepoolTotal alleles in a random mating population; Primary (same sp.) / Secondary (related sp.) / Tertiary (distant sp., embryo rescue needed)
EpistasisOne gene affects expression of another non-allelic gene; ratios: 12:3:1 (dominant), 9:3:4 (recessive), 15:1 (duplicate), 9:7 (complementary), 13:3 (inhibitory)
PleiotropismOne gene affects more than one character (e.g., seed coat colour gene in peas)
Silent MutationNucleotide change with no change in amino acid due to degeneracy of genetic code
TautomerismMolecule exists in more than one chemical form; tautomeric shifts in DNA bases → transition mutations
Pseudo allelesGovern different expressions of same character; show cis-trans position effect; can be separated by crossing over
Heterosis (Hybrid Vigour)F1 superiority over parents; coined by G.H. Shull (1914); types: relative heterosis, heterobeltiosis, standard/economic heterosis
HeterobeltiosisHeterosis measured over the better/superior parent; most useful for practical breeding
Luxuriance (Pseudo Heterosis)F1 superiority in vegetative growth only — NOT in yield or adaptation
Top CrossInbred line × open-pollinated variety; tests GCA; proposed by Davis (1927)
InbreedingMating between related individuals; selfing = highest degree; increases homozygosity 50% per generation
Inbreeding DepressionReduction in vigour/fertility in cross-pollinated species upon selfing; highest in Carrot, Alfalfa, Lucerne
AcclimatizationAdaptation of an introduced variety to a new environment
Synthetic VarietyAll combinations of selected lines + open pollination; lasts 4-5 years without yield drop; proposed by Sewall Wright
Composite VarietySeeds of several outstanding lines mixed + open pollination; broader genetic base than synthetic
Land racesTraditional cultivars developed by natural/human selection over centuries; good source of variability
ChimeraIndividual with two different genotypes in different body parts; arises from somatic mutation; e.g., Jaganath rice (gamma-ray mutant from T-141)
AmphidiploidAllopolyploid with two complete parental genomes; behaves as diploid (bivalent pairing); used as bridge species; e.g., Raphanobrassica (2n=36) by Karpechenko (1927)
IdeotypeBiological model expected to perform predictably in a defined environment; concept by Donald (1968)
CytogeneticsStudy of chromosomes and their effects on character development
GenomeComplete set of chromosomes of a diploid species (one haploid set)
EuphenicsControl of hereditary diseases via environmental/medical intervention; term by Lederberg
Multiple AllelesMany alleles of a single gene in a population; e.g., ABO blood group (I^A, I^B, i)
Cytoplasmic InheritanceCharacters controlled by plasmagenes (mitochondria, chloroplasts); maternal inheritance; e.g., male sterility in maize
First artificial plant hybridThomas Fairchild (1717) — Sweet William × Carnation = Fairchild’s Mule
Back CrossF1 × one of its parents; Recurrent parent (repeated backcross) vs Donor parent (gene donor)
Test CrossF1 × homozygous recessive parent; determines genotype of dominant phenotype; by Mendel
Pure Line TheoryBy Johannsen (1903); selection within a pure line is ineffective
Law of Homologous SeriesBy N.I. Vavilov (1920); genetically related species show parallel variation
Mutation TheoryBy Hugo de Vries (1901); based on Oenothera lamarckiana; what he saw were chromosomal aberrations
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