Lesson
04 of 9
Translate

🐞Biological Control — Parasitoids, Predators, and Microbial Agents

Natural enemies used in Indian agriculture — parasitoid-crop-pest associations, key predators, microbial biocontrol agents (Bt, NPV, Beauveria), insect traps, and cocoon types with exam mnemonics

Nature’s Own Pest Control

The previous lesson covered cultural, physical, and mechanical control — methods that modify the environment or physically remove pests. This lesson introduces the fourth IPM component: biological control, which harnesses living organisms to suppress pest populations.

In 1888, California’s citrus industry was being destroyed by the cottony cushion scale — an invasive pest from Australia. Instead of chemicals, scientists introduced the Vedalia beetle (Rodolia cardinalis), a natural predator from Australia. Within two years, the scale was under complete control. This landmark success proved that nature’s own enemies can be more effective than any chemical — and it launched the modern science of biological control.

Indian agriculture uses biological control extensively today. Trichogramma parasitoid wasps are released on millions of hectares of sugarcane and rice. Chrysoperla lacewings protect cotton from aphids. Understanding these natural enemies — who eats whom, who parasitises whom — is essential for both farming practice and competitive exams.

This lesson covers:

  1. Parasitoids — crop-pest-parasitoid associations (the most-tested table)
  2. Predators — ladybird beetles, lacewings, Cryptolaemus
  3. Microbial agents — Bt, NPV, Beauveria, Metarhizium
  4. Insect collecting traps and cocoon types

What Is Biological Control?

Biological control is the use of natural enemies — parasitoids, predators, and pathogens — to suppress pest populations below the Economic Injury Level (EIL).

Three categories of natural enemies are used:

  1. Parasitoids — insects whose larvae develop inside or on a single host, killing it
  2. Predators — organisms that kill and consume multiple prey during their lifetime
  3. Microbial agents — bacteria, viruses, and fungi that cause disease in insects

IMPORTANT

The parasitoid tables below are among the most frequently asked topics in IBPS-AFO, NABARD, and ICAR-JRF exams. Focus on memorising the crop-pest-parasitoid associations.


1. Parasitoids

A parasitoid is an insect whose immature stage develops on or within a single host insect, eventually killing it. The adult parasitoid is free-living and does not feed on the host.

Key difference from parasites: A parasite weakens but does not kill its host. A parasitoid always kills its host.

Important Parasitoids in Indian Agriculture

CropPestParasitoidExam Note
CottonHelicoverpa armigera, Pectinophora gossypiellaBracon hebetorLarval parasitoid
SugarcaneTop shoot borerIsotima javensis
MaizeStem borer (Chilo partellus)Apanteles flavis
RiceGall midge (Orselia oryzae)Platygaster oryzae
CitrusMealy bugLeptomastix dactylopiiAsked in RRB SO-2019
TobaccoSpodoptera lituraTelenomus remusEgg parasitoid
SugarcaneStem borer and top borerTrichogramma chilonisMost widely used egg parasitoid
RiceStem borerTrichogramma japonicumSpecific to rice
SugarcanePyrilla perpusellaEpiricania melanoleucaLarval parasitoid
AppleWoolly aphidAphelinus maliClassical biocontrol success
SugarcaneShoot borerSturmiopsis inferens
AppleSan Jose scaleEncarsia permiciosi
Sugarcane & CoconutShoot borer & Black Headed Caterpillar (BHC)Tetrastichus israelii
CoconutBlack headed caterpillarTrichospilus pupivoraPupal parasitoid
CottonSpotted bollworm (Earias sp.)Chilonus blackburnii
Cotton & SugarcanePink bollworm & top shoot borerBracon kirkpotricki
Sugarcane & CoconutTop shoot borer & BHCBracon bravicornis
PotatoTuber mothCopidosoma kochleri
Gram, CottonPod borer (Helicoverpa armigera)Campoletis chloridae
CottonBollworms (H. armigera, Earias, P. gossypiella)Trichogramma japonicum
Greenhouse cropsWhiteflyEncarsia formosaKey greenhouse biocontrol agent

Parasitoid Classification by Stage Attacked

ParasitoidFamilyTarget PestStage Attacked
Trichogramma sp.TrichogrammatidaeLepidopteran pests (bollworms, stem borers)Egg parasitoid
Apanteles glomeratusBraconidaeCabbage butterflyLarval parasitoid
Cotesia plutellaeBraconidaeDBM (Plutella xylostella)Larval parasitoid
Gryon sp.ScelionidaePainted bugEgg parasitoid
Alophora sp.TachinidaePainted bugAdult parasitoid

IMPORTANT

Trichogramma is the egg parasitoid of Lepidopterans. It is mass-reared on Corcyra cephalonica (Rice moth) as a factitious host — a host that is not the natural host but is used for commercial production. Corcyra eggs are easy to produce in large quantities in the laboratory.

TIP

“Tricho-Egg-Lepi”Trichogramma parasitises the Egg stage of Lepidopterans. If an exam asks about egg parasitoids of Lepidoptera, the answer is always Trichogramma.

TIP

Trichogramma memory aid: Trichogramma is the most widely used egg parasitoid in India.

  • T. chilonisChilonis for sugarcane (Cane)
  • T. japonicumJaponicum for rice (paddy, Japonica rice)

2. Predators

Predators kill and consume multiple prey during their lifetime. Unlike parasitoids, they are usually larger than their prey and do not develop inside the host.

Important Predators in Indian Agriculture

PredatorCommon NamePrey / Target PestAgricultural Use
CoccinellidsLadybird beetlesAphids, scale insects, mealy bugsNatural populations conserved; some species mass-reared
Chrysoperla carneaGreen lacewingAphids, whiteflies, bollworm eggsMass-released in cotton and vegetable fields
Cryptolaemus montrouzieriAustralian ladybird beetleMealy bugsMass-released in citrus, grape, and mango orchards
SpidersVarious speciesSmall insects in rice and cottonNatural biocontrol; conserved by reducing broad-spectrum sprays
DragonfliesMosquitoes, small flying insectsNatural predators in wetland agroecosystems
Praying mantisVarious insectsGeneralist predator in gardens and orchards
Eublemma amabilisWhite moth (Noctuidae)Lac insect (Kerria lacca)Predator of lac insects; most destructive lac pest
Holococera pulvereaBlack/grey moth (Blastobasidae)Lac insect (Kerria lacca)Predator of lac insects; second major lac enemy

NOTE

Exam favourite: Cryptolaemus montrouzieri (Australian ladybird beetle) is the key predator of mealy bugs. It is mass-reared and released in citrus, grape, and mango orchards across India.

Parasitoids vs Predators — A Comparison

FeatureParasitoidPredator
Size relative to host/preySmaller than or equal to hostLarger than prey
Number of hosts/prey consumedOne (immature stage kills one host)Many prey consumed in a lifetime
DevelopmentImmature develops on/in hostNo development on prey
Adult feedingFree-living; feeds on nectar/pollenFeeds on prey throughout life
SpecificityUsually host-specificOften generalist
ExampleTrichogramma on sugarcane borer eggsLadybird beetle eating aphids

3. Microbial Biocontrol Agents

Microbial agents are bacteria, viruses, and fungi that cause disease in pest insects. They offer the advantage of being highly specific — they kill the target pest without harming beneficial insects or humans.

BioagentTypeTarget PestKey Fact
Bacillus thuringiensis (Bt)BacteriumLepidopteran larvae (caterpillars)Produces Cry toxin; basis for Bt crops (Bt cotton, Bt brinjal)
NPV (Nuclear Polyhedrosis Virus)VirusHelicoverpa armigera, Spodoptera lituraHighly specific; each NPV strain attacks only one pest species
Beauveria bassianaFungusVarious insects (coffee berry borer, whitefly)White muscardine disease; works best in humid conditions
Metarhizium anisopliaeFungusWhite grubs, termitesGreen muscardine disease; applied as soil treatment
TrichodermaFungusSoil-borne plant pathogens (NOT insects)Biocontrol of Fusarium, Rhizoctonia; often confused with insect biocontrol

TIP

Do not confuse Trichoderma with insect biocontrol. Trichoderma controls soil-borne plant diseases (fungi), not insects. If an exam lists it alongside Bt and NPV, it is testing whether you know the difference.

Stomach vs Contact Mode of Action in Bioagents

AgentMode of Action
Bt (Bacillus thuringiensis) and NPVStomach poison — must be ingested; Bt toxin dissolves in the alkaline gut of Lepidoptera and becomes toxic
Entomopathogenic Fungi (Beauveria, Metarhizium) and NematodesContact action — infect through the cuticle; do not need to be ingested

NPV and Silkworm Disease

  • NPV causes Grasserie disease (jaundice) of silkworm (Bombyx mori)
  • Infected larvae become swollen, yellowish, and their body contents liquefy
  • Larvae hang inverted from branches/rearing trays in the characteristic symptom called Caterpillar wilt

Insect Collecting Traps and Equipment

Field monitoring requires the right trap for the right insect. This table links specific pests to their recommended collection method.

Target InsectTrap / EquipmentPrinciple
BPH (Brown Planthopper)Water trapBPH drops into water when disturbed
GrasshopperHand net (sweep net)Sweep through vegetation to catch hopping insects
WhitefliesSticky trap, suction trapYellow colour attracts whiteflies; suction pulls them in
Nocturnal mothsLight trapPositive phototaxis attracts moths at night
Specific moth speciesPheromone trap (sex lure)Synthetic pheromone attracts males of target species
House flyFood lure (molasses)Attracted to fermenting sugar

Types of Cocoons

Different insect orders and species construct different cocoon types during pupation. This classification is a niche but regularly tested topic.

Cocoon TypeMaterial UsedExample InsectExam Detail
Silken cocoonSilkSilkworm (Bombyx mori)Commercially harvested for silk
Earthen cocoonSoil + salivaGram pod borer (Helicoverpa)Pupates in soil
Hairy cocoonBody hairsWoolly bears (Arctiidae)Hairs from larval body
Frassy cocoonFrass (excrement) + salivaCoconut black headed caterpillarUnique material
Fibrous cocoonPlant fibresRed palm weevilFound inside palm trunk
PupariumHardened last larval skinHouse fly, fruit flyNOT a true cocoon

IMPORTANT

Puparium is NOT a true cocoon. It is the hardened last larval skin that serves as a protective case for the pupa. It is characteristic of Diptera (flies). This distinction is a frequent exam question.


When to Use Which Biocontrol Agent?

Quick decision guide for field officers:

Pest to ControlBest Biocontrol AgentTypeWhen to Release/ApplyDose
Bollworm/stem borer eggsTrichogramma spp.Egg parasitoidAt egg-laying peak (use pheromone trap to monitor)50,000-1,00,000/ha
Caterpillars (Lepidoptera)Bacillus thuringiensis (Bt)BacteriumEarly larval instars (L1-L2); evening spray1-2 kg/ha
Helicoverpa larvaeHaNPVVirusEarly instars; evening application (UV-sensitive)250-500 LE/ha
MealybugCryptolaemus montrouzieriPredator beetleWhen colonies visible10 beetles/tree
AphidsChrysoperla carnea (lacewing)PredatorWhen aphid colonies appear50,000 eggs/ha
Pyrilla (sugarcane)Epiricania melanoleucaParasitoidRelease cocoons on infested leaves4000-5000 cocoons/ha
White grubs in soilMetarhizium anisopliaeFungusSoil application before planting5 kg/ha mixed with FYM
Fruit flySterile Insect Technique (SIT)PhysicalContinuous area-wide releaseGovt. programme

Critical timing rule: Biocontrol agents must be released before pest population explodes. Once pest is at epidemic level, biocontrol alone cannot save the crop — you’ll need to integrate with other methods.


Exam Tips and Mnemonics

  1. Vedalia beetle story — the first major biological control success (1888, California, cottony cushion scale). Know the pest, predator, and country.
  2. Trichogramma = egg parasitoid. If the question says “egg parasitoid,” think Trichogramma first.
  3. Cryptolaemus = mealy bug predator. The name sounds like “crypto-lame-us” — mealy bugs are cryptic (hidden) and lame (slow-moving).
  4. NPV is virus, Bt is bacterium, Beauveria is fungus — know the type of each microbial agent.
  5. Puparium belongs to Diptera. All other cocoon types belong to Lepidoptera or Coleoptera.

Summary Table

CategoryKey AgentsTarget PestsExam Priority
Egg parasitoidsTrichogramma chilonis, T. japonicum, Telenomus remusSugarcane/rice borers, SpodopteraVery high
Larval parasitoidsBracon hebetor, Apanteles flavis, Epiricania melanoleucaCotton bollworm, maize borer, pyrillaHigh
PredatorsLadybird beetles, Chrysoperla, CryptolaemusAphids, whiteflies, mealy bugsHigh
BacterialBacillus thuringiensis (Bt)Lepidopteran larvaeVery high
ViralNPV (Nuclear Polyhedrosis Virus)Helicoverpa, SpodopteraHigh
FungalBeauveria bassiana, Metarhizium anisopliaeVarious insects, white grubsModerate

Summary Cheat Sheet

Concept / TopicKey Details
Vedalia beetleRodolia cardinalis; 1888 California; controlled cottony cushion scale — first major biocontrol success
Parasitoid vs ParasiteParasitoid always kills host; parasite weakens but does not kill
Trichogramma chilonisMost widely used egg parasitoid in India — sugarcane borers
Trichogramma japonicumEgg parasitoid specific to rice stem borer
Cryptolaemus montrouzieriAustralian ladybird beetle — key predator of mealy bugs in citrus, grape, mango
Chrysoperla carneaGreen lacewing — predator of aphids, whiteflies, bollworm eggs in cotton
Bt (Bacillus thuringiensis)Bacterium; Cry toxin; targets Lepidopteran larvae; basis for Bt crops
NPVNuclear Polyhedrosis Virus; highly specific — each strain attacks one pest species
Beauveria bassianaFungus; white muscardine disease; works best in humid conditions
Metarhizium anisopliaeFungus; green muscardine disease; soil treatment for white grubs, termites
TrichodermaControls soil-borne plant diseases (fungi), NOT insects — common exam confusion
PupariumNOT a true cocoon; hardened last larval skin; characteristic of Diptera (flies)

TIP

Next: Lesson 05 covers Chemical Control — insecticide generations, formulations, toxicology, and the WHO hazard classification that appears on every pesticide label.

🔐

Pro Content Locked

Upgrade to Pro to access this lesson and all other premium content.

Pro Popular
199 /mo

₹2388 billed yearly

  • All Agriculture & Banking Courses
  • AI Lesson Questions (100/day)
  • AI Doubt Solver (50/day)
  • Glows & Grows Feedback (30/day)
  • AI Section Quiz (20/day)
  • 22-Language Translation (30/day)
  • Recall Questions (20/day)
  • AI Quiz (15/day)
  • AI Quiz Paper Analysis
  • AI Step-by-Step Explanations
  • Spaced Repetition Recall (FSRS)
  • AI Tutor
  • Immersive Text Questions
  • Audio Lessons — Hindi & English
  • Mock Tests & Previous Year Papers
  • Summary & Mind Maps
  • XP, Levels, Leaderboard & Badges
  • Generate New Classrooms
  • Voice AI Teacher (AgriDots Live)
  • AI Revision Assistant
  • Knowledge Gap Analysis
  • Interactive Revision (LangGraph)

🔒 Secure via Razorpay · Cancel anytime · No hidden fees

Lesson Doubts

Ask questions, get expert answers

Lesson Doubts is a Pro feature.Upgrade