🐞Biological Control — Parasitoids, Predators, and Microbial Agents
Natural enemies used in Indian agriculture — parasitoid-crop-pest associations, key predators, microbial biocontrol agents (Bt, NPV, Beauveria), insect traps, and cocoon types with exam mnemonics
Nature’s Own Pest Control
The previous lesson covered cultural, physical, and mechanical control — methods that modify the environment or physically remove pests. This lesson introduces the fourth IPM component: biological control, which harnesses living organisms to suppress pest populations.
In 1888, California’s citrus industry was being destroyed by the cottony cushion scale — an invasive pest from Australia. Instead of chemicals, scientists introduced the Vedalia beetle (Rodolia cardinalis), a natural predator from Australia. Within two years, the scale was under complete control. This landmark success proved that nature’s own enemies can be more effective than any chemical — and it launched the modern science of biological control.
Indian agriculture uses biological control extensively today. Trichogramma parasitoid wasps are released on millions of hectares of sugarcane and rice. Chrysoperla lacewings protect cotton from aphids. Understanding these natural enemies — who eats whom, who parasitises whom — is essential for both farming practice and competitive exams.
This lesson covers:
- Parasitoids — crop-pest-parasitoid associations (the most-tested table)
- Predators — ladybird beetles, lacewings, Cryptolaemus
- Microbial agents — Bt, NPV, Beauveria, Metarhizium
- Insect collecting traps and cocoon types
What Is Biological Control?
Biological control is the use of natural enemies — parasitoids, predators, and pathogens — to suppress pest populations below the Economic Injury Level (EIL).
Three categories of natural enemies are used:
- Parasitoids — insects whose larvae develop inside or on a single host, killing it
- Predators — organisms that kill and consume multiple prey during their lifetime
- Microbial agents — bacteria, viruses, and fungi that cause disease in insects
IMPORTANT
The parasitoid tables below are among the most frequently asked topics in IBPS-AFO, NABARD, and ICAR-JRF exams. Focus on memorising the crop-pest-parasitoid associations.
1. Parasitoids
A parasitoid is an insect whose immature stage develops on or within a single host insect, eventually killing it. The adult parasitoid is free-living and does not feed on the host.
Key difference from parasites: A parasite weakens but does not kill its host. A parasitoid always kills its host.
Important Parasitoids in Indian Agriculture
| Crop | Pest | Parasitoid | Exam Note |
|---|---|---|---|
| Cotton | Helicoverpa armigera, Pectinophora gossypiella | Bracon hebetor | Larval parasitoid |
| Sugarcane | Top shoot borer | Isotima javensis | |
| Maize | Stem borer (Chilo partellus) | Apanteles flavis | |
| Rice | Gall midge (Orselia oryzae) | Platygaster oryzae | |
| Citrus | Mealy bug | Leptomastix dactylopii | Asked in RRB SO-2019 |
| Tobacco | Spodoptera litura | Telenomus remus | Egg parasitoid |
| Sugarcane | Stem borer and top borer | Trichogramma chilonis | Most widely used egg parasitoid |
| Rice | Stem borer | Trichogramma japonicum | Specific to rice |
| Sugarcane | Pyrilla perpusella | Epiricania melanoleuca | Larval parasitoid |
| Apple | Woolly aphid | Aphelinus mali | Classical biocontrol success |
| Sugarcane | Shoot borer | Sturmiopsis inferens | |
| Apple | San Jose scale | Encarsia permiciosi | |
| Sugarcane & Coconut | Shoot borer & Black Headed Caterpillar (BHC) | Tetrastichus israelii | |
| Coconut | Black headed caterpillar | Trichospilus pupivora | Pupal parasitoid |
| Cotton | Spotted bollworm (Earias sp.) | Chilonus blackburnii | |
| Cotton & Sugarcane | Pink bollworm & top shoot borer | Bracon kirkpotricki | |
| Sugarcane & Coconut | Top shoot borer & BHC | Bracon bravicornis | |
| Potato | Tuber moth | Copidosoma kochleri | |
| Gram, Cotton | Pod borer (Helicoverpa armigera) | Campoletis chloridae | |
| Cotton | Bollworms (H. armigera, Earias, P. gossypiella) | Trichogramma japonicum | |
| Greenhouse crops | Whitefly | Encarsia formosa | Key greenhouse biocontrol agent |
Parasitoid Classification by Stage Attacked
| Parasitoid | Family | Target Pest | Stage Attacked |
|---|---|---|---|
| Trichogramma sp. | Trichogrammatidae | Lepidopteran pests (bollworms, stem borers) | Egg parasitoid |
| Apanteles glomeratus | Braconidae | Cabbage butterfly | Larval parasitoid |
| Cotesia plutellae | Braconidae | DBM (Plutella xylostella) | Larval parasitoid |
| Gryon sp. | Scelionidae | Painted bug | Egg parasitoid |
| Alophora sp. | Tachinidae | Painted bug | Adult parasitoid |
IMPORTANT
Trichogramma is the egg parasitoid of Lepidopterans. It is mass-reared on Corcyra cephalonica (Rice moth) as a factitious host — a host that is not the natural host but is used for commercial production. Corcyra eggs are easy to produce in large quantities in the laboratory.
TIP
“Tricho-Egg-Lepi” — Trichogramma parasitises the Egg stage of Lepidopterans. If an exam asks about egg parasitoids of Lepidoptera, the answer is always Trichogramma.
TIP
Trichogramma memory aid: Trichogramma is the most widely used egg parasitoid in India.
- T. chilonis → Chilonis for sugarcane (Cane)
- T. japonicum → Japonicum for rice (paddy, Japonica rice)
2. Predators
Predators kill and consume multiple prey during their lifetime. Unlike parasitoids, they are usually larger than their prey and do not develop inside the host.
Important Predators in Indian Agriculture
| Predator | Common Name | Prey / Target Pest | Agricultural Use |
|---|---|---|---|
| Coccinellids | Ladybird beetles | Aphids, scale insects, mealy bugs | Natural populations conserved; some species mass-reared |
| Chrysoperla carnea | Green lacewing | Aphids, whiteflies, bollworm eggs | Mass-released in cotton and vegetable fields |
| Cryptolaemus montrouzieri | Australian ladybird beetle | Mealy bugs | Mass-released in citrus, grape, and mango orchards |
| Spiders | Various species | Small insects in rice and cotton | Natural biocontrol; conserved by reducing broad-spectrum sprays |
| Dragonflies | — | Mosquitoes, small flying insects | Natural predators in wetland agroecosystems |
| Praying mantis | — | Various insects | Generalist predator in gardens and orchards |
| Eublemma amabilis | White moth (Noctuidae) | Lac insect (Kerria lacca) | Predator of lac insects; most destructive lac pest |
| Holococera pulverea | Black/grey moth (Blastobasidae) | Lac insect (Kerria lacca) | Predator of lac insects; second major lac enemy |
NOTE
Exam favourite: Cryptolaemus montrouzieri (Australian ladybird beetle) is the key predator of mealy bugs. It is mass-reared and released in citrus, grape, and mango orchards across India.
Parasitoids vs Predators — A Comparison
| Feature | Parasitoid | Predator |
|---|---|---|
| Size relative to host/prey | Smaller than or equal to host | Larger than prey |
| Number of hosts/prey consumed | One (immature stage kills one host) | Many prey consumed in a lifetime |
| Development | Immature develops on/in host | No development on prey |
| Adult feeding | Free-living; feeds on nectar/pollen | Feeds on prey throughout life |
| Specificity | Usually host-specific | Often generalist |
| Example | Trichogramma on sugarcane borer eggs | Ladybird beetle eating aphids |
3. Microbial Biocontrol Agents
Microbial agents are bacteria, viruses, and fungi that cause disease in pest insects. They offer the advantage of being highly specific — they kill the target pest without harming beneficial insects or humans.
| Bioagent | Type | Target Pest | Key Fact |
|---|---|---|---|
| Bacillus thuringiensis (Bt) | Bacterium | Lepidopteran larvae (caterpillars) | Produces Cry toxin; basis for Bt crops (Bt cotton, Bt brinjal) |
| NPV (Nuclear Polyhedrosis Virus) | Virus | Helicoverpa armigera, Spodoptera litura | Highly specific; each NPV strain attacks only one pest species |
| Beauveria bassiana | Fungus | Various insects (coffee berry borer, whitefly) | White muscardine disease; works best in humid conditions |
| Metarhizium anisopliae | Fungus | White grubs, termites | Green muscardine disease; applied as soil treatment |
| Trichoderma | Fungus | Soil-borne plant pathogens (NOT insects) | Biocontrol of Fusarium, Rhizoctonia; often confused with insect biocontrol |
TIP
Do not confuse Trichoderma with insect biocontrol. Trichoderma controls soil-borne plant diseases (fungi), not insects. If an exam lists it alongside Bt and NPV, it is testing whether you know the difference.
Stomach vs Contact Mode of Action in Bioagents
| Agent | Mode of Action |
|---|---|
| Bt (Bacillus thuringiensis) and NPV | Stomach poison — must be ingested; Bt toxin dissolves in the alkaline gut of Lepidoptera and becomes toxic |
| Entomopathogenic Fungi (Beauveria, Metarhizium) and Nematodes | Contact action — infect through the cuticle; do not need to be ingested |
NPV and Silkworm Disease
- NPV causes Grasserie disease (jaundice) of silkworm (Bombyx mori)
- Infected larvae become swollen, yellowish, and their body contents liquefy
- Larvae hang inverted from branches/rearing trays in the characteristic symptom called Caterpillar wilt
Insect Collecting Traps and Equipment
Field monitoring requires the right trap for the right insect. This table links specific pests to their recommended collection method.
| Target Insect | Trap / Equipment | Principle |
|---|---|---|
| BPH (Brown Planthopper) | Water trap | BPH drops into water when disturbed |
| Grasshopper | Hand net (sweep net) | Sweep through vegetation to catch hopping insects |
| Whiteflies | Sticky trap, suction trap | Yellow colour attracts whiteflies; suction pulls them in |
| Nocturnal moths | Light trap | Positive phototaxis attracts moths at night |
| Specific moth species | Pheromone trap (sex lure) | Synthetic pheromone attracts males of target species |
| House fly | Food lure (molasses) | Attracted to fermenting sugar |
Types of Cocoons
Different insect orders and species construct different cocoon types during pupation. This classification is a niche but regularly tested topic.
| Cocoon Type | Material Used | Example Insect | Exam Detail |
|---|---|---|---|
| Silken cocoon | Silk | Silkworm (Bombyx mori) | Commercially harvested for silk |
| Earthen cocoon | Soil + saliva | Gram pod borer (Helicoverpa) | Pupates in soil |
| Hairy cocoon | Body hairs | Woolly bears (Arctiidae) | Hairs from larval body |
| Frassy cocoon | Frass (excrement) + saliva | Coconut black headed caterpillar | Unique material |
| Fibrous cocoon | Plant fibres | Red palm weevil | Found inside palm trunk |
| Puparium | Hardened last larval skin | House fly, fruit fly | NOT a true cocoon |
IMPORTANT
Puparium is NOT a true cocoon. It is the hardened last larval skin that serves as a protective case for the pupa. It is characteristic of Diptera (flies). This distinction is a frequent exam question.
When to Use Which Biocontrol Agent?
Quick decision guide for field officers:
| Pest to Control | Best Biocontrol Agent | Type | When to Release/Apply | Dose |
|---|---|---|---|---|
| Bollworm/stem borer eggs | Trichogramma spp. | Egg parasitoid | At egg-laying peak (use pheromone trap to monitor) | 50,000-1,00,000/ha |
| Caterpillars (Lepidoptera) | Bacillus thuringiensis (Bt) | Bacterium | Early larval instars (L1-L2); evening spray | 1-2 kg/ha |
| Helicoverpa larvae | HaNPV | Virus | Early instars; evening application (UV-sensitive) | 250-500 LE/ha |
| Mealybug | Cryptolaemus montrouzieri | Predator beetle | When colonies visible | 10 beetles/tree |
| Aphids | Chrysoperla carnea (lacewing) | Predator | When aphid colonies appear | 50,000 eggs/ha |
| Pyrilla (sugarcane) | Epiricania melanoleuca | Parasitoid | Release cocoons on infested leaves | 4000-5000 cocoons/ha |
| White grubs in soil | Metarhizium anisopliae | Fungus | Soil application before planting | 5 kg/ha mixed with FYM |
| Fruit fly | Sterile Insect Technique (SIT) | Physical | Continuous area-wide release | Govt. programme |
Critical timing rule: Biocontrol agents must be released before pest population explodes. Once pest is at epidemic level, biocontrol alone cannot save the crop — you’ll need to integrate with other methods.
Exam Tips and Mnemonics
- Vedalia beetle story — the first major biological control success (1888, California, cottony cushion scale). Know the pest, predator, and country.
- Trichogramma = egg parasitoid. If the question says “egg parasitoid,” think Trichogramma first.
- Cryptolaemus = mealy bug predator. The name sounds like “crypto-lame-us” — mealy bugs are cryptic (hidden) and lame (slow-moving).
- NPV is virus, Bt is bacterium, Beauveria is fungus — know the type of each microbial agent.
- Puparium belongs to Diptera. All other cocoon types belong to Lepidoptera or Coleoptera.
Summary Table
| Category | Key Agents | Target Pests | Exam Priority |
|---|---|---|---|
| Egg parasitoids | Trichogramma chilonis, T. japonicum, Telenomus remus | Sugarcane/rice borers, Spodoptera | Very high |
| Larval parasitoids | Bracon hebetor, Apanteles flavis, Epiricania melanoleuca | Cotton bollworm, maize borer, pyrilla | High |
| Predators | Ladybird beetles, Chrysoperla, Cryptolaemus | Aphids, whiteflies, mealy bugs | High |
| Bacterial | Bacillus thuringiensis (Bt) | Lepidopteran larvae | Very high |
| Viral | NPV (Nuclear Polyhedrosis Virus) | Helicoverpa, Spodoptera | High |
| Fungal | Beauveria bassiana, Metarhizium anisopliae | Various insects, white grubs | Moderate |
Summary Cheat Sheet
| Concept / Topic | Key Details |
|---|---|
| Vedalia beetle | Rodolia cardinalis; 1888 California; controlled cottony cushion scale — first major biocontrol success |
| Parasitoid vs Parasite | Parasitoid always kills host; parasite weakens but does not kill |
| Trichogramma chilonis | Most widely used egg parasitoid in India — sugarcane borers |
| Trichogramma japonicum | Egg parasitoid specific to rice stem borer |
| Cryptolaemus montrouzieri | Australian ladybird beetle — key predator of mealy bugs in citrus, grape, mango |
| Chrysoperla carnea | Green lacewing — predator of aphids, whiteflies, bollworm eggs in cotton |
| Bt (Bacillus thuringiensis) | Bacterium; Cry toxin; targets Lepidopteran larvae; basis for Bt crops |
| NPV | Nuclear Polyhedrosis Virus; highly specific — each strain attacks one pest species |
| Beauveria bassiana | Fungus; white muscardine disease; works best in humid conditions |
| Metarhizium anisopliae | Fungus; green muscardine disease; soil treatment for white grubs, termites |
| Trichoderma | Controls soil-borne plant diseases (fungi), NOT insects — common exam confusion |
| Puparium | NOT a true cocoon; hardened last larval skin; characteristic of Diptera (flies) |
TIP
Next: Lesson 05 covers Chemical Control — insecticide generations, formulations, toxicology, and the WHO hazard classification that appears on every pesticide label.
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Nature’s Own Pest Control
The previous lesson covered cultural, physical, and mechanical control — methods that modify the environment or physically remove pests. This lesson introduces the fourth IPM component: biological control, which harnesses living organisms to suppress pest populations.
In 1888, California’s citrus industry was being destroyed by the cottony cushion scale — an invasive pest from Australia. Instead of chemicals, scientists introduced the Vedalia beetle (Rodolia cardinalis), a natural predator from Australia. Within two years, the scale was under complete control. This landmark success proved that nature’s own enemies can be more effective than any chemical — and it launched the modern science of biological control.
Indian agriculture uses biological control extensively today. Trichogramma parasitoid wasps are released on millions of hectares of sugarcane and rice. Chrysoperla lacewings protect cotton from aphids. Understanding these natural enemies — who eats whom, who parasitises whom — is essential for both farming practice and competitive exams.
This lesson covers:
- Parasitoids — crop-pest-parasitoid associations (the most-tested table)
- Predators — ladybird beetles, lacewings, Cryptolaemus
- Microbial agents — Bt, NPV, Beauveria, Metarhizium
- Insect collecting traps and cocoon types
What Is Biological Control?
Biological control is the use of natural enemies — parasitoids, predators, and pathogens — to suppress pest populations below the Economic Injury Level (EIL).
Three categories of natural enemies are used:
- Parasitoids — insects whose larvae develop inside or on a single host, killing it
- Predators — organisms that kill and consume multiple prey during their lifetime
- Microbial agents — bacteria, viruses, and fungi that cause disease in insects
IMPORTANT
The parasitoid tables below are among the most frequently asked topics in IBPS-AFO, NABARD, and ICAR-JRF exams. Focus on memorising the crop-pest-parasitoid associations.
1. Parasitoids
A parasitoid is an insect whose immature stage develops on or within a single host insect, eventually killing it. The adult parasitoid is free-living and does not feed on the host.
Key difference from parasites: A parasite weakens but does not kill its host. A parasitoid always kills its host.
Important Parasitoids in Indian Agriculture
| Crop | Pest | Parasitoid | Exam Note |
|---|---|---|---|
| Cotton | Helicoverpa armigera, Pectinophora gossypiella | Bracon hebetor | Larval parasitoid |
| Sugarcane | Top shoot borer | Isotima javensis | |
| Maize | Stem borer (Chilo partellus) | Apanteles flavis | |
| Rice | Gall midge (Orselia oryzae) | Platygaster oryzae | |
| Citrus | Mealy bug | Leptomastix dactylopii | Asked in RRB SO-2019 |
| Tobacco | Spodoptera litura | Telenomus remus | Egg parasitoid |
| Sugarcane | Stem borer and top borer | Trichogramma chilonis | Most widely used egg parasitoid |
| Rice | Stem borer | Trichogramma japonicum | Specific to rice |
| Sugarcane | Pyrilla perpusella | Epiricania melanoleuca | Larval parasitoid |
| Apple | Woolly aphid | Aphelinus mali | Classical biocontrol success |
| Sugarcane | Shoot borer | Sturmiopsis inferens | |
| Apple | San Jose scale | Encarsia permiciosi | |
| Sugarcane & Coconut | Shoot borer & Black Headed Caterpillar (BHC) | Tetrastichus israelii | |
| Coconut | Black headed caterpillar | Trichospilus pupivora | Pupal parasitoid |
| Cotton | Spotted bollworm (Earias sp.) | Chilonus blackburnii | |
| Cotton & Sugarcane | Pink bollworm & top shoot borer | Bracon kirkpotricki | |
| Sugarcane & Coconut | Top shoot borer & BHC | Bracon bravicornis | |
| Potato | Tuber moth | Copidosoma kochleri | |
| Gram, Cotton | Pod borer (Helicoverpa armigera) | Campoletis chloridae | |
| Cotton | Bollworms (H. armigera, Earias, P. gossypiella) | Trichogramma japonicum | |
| Greenhouse crops | Whitefly | Encarsia formosa | Key greenhouse biocontrol agent |
Parasitoid Classification by Stage Attacked
| Parasitoid | Family | Target Pest | Stage Attacked |
|---|---|---|---|
| Trichogramma sp. | Trichogrammatidae | Lepidopteran pests (bollworms, stem borers) | Egg parasitoid |
| Apanteles glomeratus | Braconidae | Cabbage butterfly | Larval parasitoid |
| Cotesia plutellae | Braconidae | DBM (Plutella xylostella) | Larval parasitoid |
| Gryon sp. | Scelionidae | Painted bug | Egg parasitoid |
| Alophora sp. | Tachinidae | Painted bug | Adult parasitoid |
IMPORTANT
Trichogramma is the egg parasitoid of Lepidopterans. It is mass-reared on Corcyra cephalonica (Rice moth) as a factitious host — a host that is not the natural host but is used for commercial production. Corcyra eggs are easy to produce in large quantities in the laboratory.
TIP
“Tricho-Egg-Lepi” — Trichogramma parasitises the Egg stage of Lepidopterans. If an exam asks about egg parasitoids of Lepidoptera, the answer is always Trichogramma.
TIP
Trichogramma memory aid: Trichogramma is the most widely used egg parasitoid in India.
- T. chilonis → Chilonis for sugarcane (Cane)
- T. japonicum → Japonicum for rice (paddy, Japonica rice)
2. Predators
Predators kill and consume multiple prey during their lifetime. Unlike parasitoids, they are usually larger than their prey and do not develop inside the host.
Important Predators in Indian Agriculture
| Predator | Common Name | Prey / Target Pest | Agricultural Use |
|---|---|---|---|
| Coccinellids | Ladybird beetles | Aphids, scale insects, mealy bugs | Natural populations conserved; some species mass-reared |
| Chrysoperla carnea | Green lacewing | Aphids, whiteflies, bollworm eggs | Mass-released in cotton and vegetable fields |
| Cryptolaemus montrouzieri | Australian ladybird beetle | Mealy bugs | Mass-released in citrus, grape, and mango orchards |
| Spiders | Various species | Small insects in rice and cotton | Natural biocontrol; conserved by reducing broad-spectrum sprays |
| Dragonflies | — | Mosquitoes, small flying insects | Natural predators in wetland agroecosystems |
| Praying mantis | — | Various insects | Generalist predator in gardens and orchards |
| Eublemma amabilis | White moth (Noctuidae) | Lac insect (Kerria lacca) | Predator of lac insects; most destructive lac pest |
| Holococera pulverea | Black/grey moth (Blastobasidae) | Lac insect (Kerria lacca) | Predator of lac insects; second major lac enemy |
NOTE
Exam favourite: Cryptolaemus montrouzieri (Australian ladybird beetle) is the key predator of mealy bugs. It is mass-reared and released in citrus, grape, and mango orchards across India.
Parasitoids vs Predators — A Comparison
| Feature | Parasitoid | Predator |
|---|---|---|
| Size relative to host/prey | Smaller than or equal to host | Larger than prey |
| Number of hosts/prey consumed | One (immature stage kills one host) | Many prey consumed in a lifetime |
| Development | Immature develops on/in host | No development on prey |
| Adult feeding | Free-living; feeds on nectar/pollen | Feeds on prey throughout life |
| Specificity | Usually host-specific | Often generalist |
| Example | Trichogramma on sugarcane borer eggs | Ladybird beetle eating aphids |
3. Microbial Biocontrol Agents
Microbial agents are bacteria, viruses, and fungi that cause disease in pest insects. They offer the advantage of being highly specific — they kill the target pest without harming beneficial insects or humans.
| Bioagent | Type | Target Pest | Key Fact |
|---|---|---|---|
| Bacillus thuringiensis (Bt) | Bacterium | Lepidopteran larvae (caterpillars) | Produces Cry toxin; basis for Bt crops (Bt cotton, Bt brinjal) |
| NPV (Nuclear Polyhedrosis Virus) | Virus | Helicoverpa armigera, Spodoptera litura | Highly specific; each NPV strain attacks only one pest species |
| Beauveria bassiana | Fungus | Various insects (coffee berry borer, whitefly) | White muscardine disease; works best in humid conditions |
| Metarhizium anisopliae | Fungus | White grubs, termites | Green muscardine disease; applied as soil treatment |
| Trichoderma | Fungus | Soil-borne plant pathogens (NOT insects) | Biocontrol of Fusarium, Rhizoctonia; often confused with insect biocontrol |
TIP
Do not confuse Trichoderma with insect biocontrol. Trichoderma controls soil-borne plant diseases (fungi), not insects. If an exam lists it alongside Bt and NPV, it is testing whether you know the difference.
Stomach vs Contact Mode of Action in Bioagents
| Agent | Mode of Action |
|---|---|
| Bt (Bacillus thuringiensis) and NPV | Stomach poison — must be ingested; Bt toxin dissolves in the alkaline gut of Lepidoptera and becomes toxic |
| Entomopathogenic Fungi (Beauveria, Metarhizium) and Nematodes | Contact action — infect through the cuticle; do not need to be ingested |
NPV and Silkworm Disease
- NPV causes Grasserie disease (jaundice) of silkworm (Bombyx mori)
- Infected larvae become swollen, yellowish, and their body contents liquefy
- Larvae hang inverted from branches/rearing trays in the characteristic symptom called Caterpillar wilt
Insect Collecting Traps and Equipment
Field monitoring requires the right trap for the right insect. This table links specific pests to their recommended collection method.
| Target Insect | Trap / Equipment | Principle |
|---|---|---|
| BPH (Brown Planthopper) | Water trap | BPH drops into water when disturbed |
| Grasshopper | Hand net (sweep net) | Sweep through vegetation to catch hopping insects |
| Whiteflies | Sticky trap, suction trap | Yellow colour attracts whiteflies; suction pulls them in |
| Nocturnal moths | Light trap | Positive phototaxis attracts moths at night |
| Specific moth species | Pheromone trap (sex lure) | Synthetic pheromone attracts males of target species |
| House fly | Food lure (molasses) | Attracted to fermenting sugar |
Types of Cocoons
Different insect orders and species construct different cocoon types during pupation. This classification is a niche but regularly tested topic.
| Cocoon Type | Material Used | Example Insect | Exam Detail |
|---|---|---|---|
| Silken cocoon | Silk | Silkworm (Bombyx mori) | Commercially harvested for silk |
| Earthen cocoon | Soil + saliva | Gram pod borer (Helicoverpa) | Pupates in soil |
| Hairy cocoon | Body hairs | Woolly bears (Arctiidae) | Hairs from larval body |
| Frassy cocoon | Frass (excrement) + saliva | Coconut black headed caterpillar | Unique material |
| Fibrous cocoon | Plant fibres | Red palm weevil | Found inside palm trunk |
| Puparium | Hardened last larval skin | House fly, fruit fly | NOT a true cocoon |
IMPORTANT
Puparium is NOT a true cocoon. It is the hardened last larval skin that serves as a protective case for the pupa. It is characteristic of Diptera (flies). This distinction is a frequent exam question.
When to Use Which Biocontrol Agent?
Quick decision guide for field officers:
| Pest to Control | Best Biocontrol Agent | Type | When to Release/Apply | Dose |
|---|---|---|---|---|
| Bollworm/stem borer eggs | Trichogramma spp. | Egg parasitoid | At egg-laying peak (use pheromone trap to monitor) | 50,000-1,00,000/ha |
| Caterpillars (Lepidoptera) | Bacillus thuringiensis (Bt) | Bacterium | Early larval instars (L1-L2); evening spray | 1-2 kg/ha |
| Helicoverpa larvae | HaNPV | Virus | Early instars; evening application (UV-sensitive) | 250-500 LE/ha |
| Mealybug | Cryptolaemus montrouzieri | Predator beetle | When colonies visible | 10 beetles/tree |
| Aphids | Chrysoperla carnea (lacewing) | Predator | When aphid colonies appear | 50,000 eggs/ha |
| Pyrilla (sugarcane) | Epiricania melanoleuca | Parasitoid | Release cocoons on infested leaves | 4000-5000 cocoons/ha |
| White grubs in soil | Metarhizium anisopliae | Fungus | Soil application before planting | 5 kg/ha mixed with FYM |
| Fruit fly | Sterile Insect Technique (SIT) | Physical | Continuous area-wide release | Govt. programme |
Critical timing rule: Biocontrol agents must be released before pest population explodes. Once pest is at epidemic level, biocontrol alone cannot save the crop — you’ll need to integrate with other methods.
Exam Tips and Mnemonics
- Vedalia beetle story — the first major biological control success (1888, California, cottony cushion scale). Know the pest, predator, and country.
- Trichogramma = egg parasitoid. If the question says “egg parasitoid,” think Trichogramma first.
- Cryptolaemus = mealy bug predator. The name sounds like “crypto-lame-us” — mealy bugs are cryptic (hidden) and lame (slow-moving).
- NPV is virus, Bt is bacterium, Beauveria is fungus — know the type of each microbial agent.
- Puparium belongs to Diptera. All other cocoon types belong to Lepidoptera or Coleoptera.
Summary Table
| Category | Key Agents | Target Pests | Exam Priority |
|---|---|---|---|
| Egg parasitoids | Trichogramma chilonis, T. japonicum, Telenomus remus | Sugarcane/rice borers, Spodoptera | Very high |
| Larval parasitoids | Bracon hebetor, Apanteles flavis, Epiricania melanoleuca | Cotton bollworm, maize borer, pyrilla | High |
| Predators | Ladybird beetles, Chrysoperla, Cryptolaemus | Aphids, whiteflies, mealy bugs | High |
| Bacterial | Bacillus thuringiensis (Bt) | Lepidopteran larvae | Very high |
| Viral | NPV (Nuclear Polyhedrosis Virus) | Helicoverpa, Spodoptera | High |
| Fungal | Beauveria bassiana, Metarhizium anisopliae | Various insects, white grubs | Moderate |
Summary Cheat Sheet
| Concept / Topic | Key Details |
|---|---|
| Vedalia beetle | Rodolia cardinalis; 1888 California; controlled cottony cushion scale — first major biocontrol success |
| Parasitoid vs Parasite | Parasitoid always kills host; parasite weakens but does not kill |
| Trichogramma chilonis | Most widely used egg parasitoid in India — sugarcane borers |
| Trichogramma japonicum | Egg parasitoid specific to rice stem borer |
| Cryptolaemus montrouzieri | Australian ladybird beetle — key predator of mealy bugs in citrus, grape, mango |
| Chrysoperla carnea | Green lacewing — predator of aphids, whiteflies, bollworm eggs in cotton |
| Bt (Bacillus thuringiensis) | Bacterium; Cry toxin; targets Lepidopteran larvae; basis for Bt crops |
| NPV | Nuclear Polyhedrosis Virus; highly specific — each strain attacks one pest species |
| Beauveria bassiana | Fungus; white muscardine disease; works best in humid conditions |
| Metarhizium anisopliae | Fungus; green muscardine disease; soil treatment for white grubs, termites |
| Trichoderma | Controls soil-borne plant diseases (fungi), NOT insects — common exam confusion |
| Puparium | NOT a true cocoon; hardened last larval skin; characteristic of Diptera (flies) |
TIP
Next: Lesson 05 covers Chemical Control — insecticide generations, formulations, toxicology, and the WHO hazard classification that appears on every pesticide label.
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